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as it were, of our digestive mechanism. It is the duty of taste to warn us against uneatable things, and to recommend to our favourable attention eatable and wholesome ones; and, on the whole, in spite of small occasional remissness, it performs this duty with creditable success.

Taste, however, is not equally distributed over the whole surface of the tongue alike. There are three distinct regions or tracts, each of which has to perform its own special office and function. The tip of the tongue is concerned mainly with pungent and acrid tastes; the middle portion is sensitive chiefly to sweets and bitters; while the back or lower portion confines itself almost entirely to the flavours of roast meats, butter, oils, and other rich or fatty substances. There are very good reasons for this subdivision of faculties in the tongue, the object being, as it were, to make each piece of food undergo three separate examinations (like 'smalls,' 'mods,' and 'greats' at Oxford), which must be successively passed before it is admitted into full participation in the human economy. The first examination, as we shall shortly see, gets rid at once of substances which would be actively and immediately destructive to the very tissues of the mouth and body; the second discriminates between poisonous and chemically harmless food-stuffs; and the third merely decides the minor question whether the particular food is likely to prove then and there wholesome or indigestible to the particular person. The sense of taste proceeds, in fact, upon the principle of gradual selection and elimination; it refuses first what is positively destructive, next what is more remotely deleterious, and finally what is only undesirable or over-luscious.

When we want to assure ourselves, by means of taste, about any unknown object—say a lump of some white stuff, which may be crystal, or glass, or alum, or borax, or quartz, or rocksalt—we put the tip of the tongue against it gingerly. If it begins to burn us, we draw it away more or less rapidly with an accompaniment in language strictly dependent upon our personal habits and manners. The test we thus occasionally apply, even in the civilised adult state, to unknown bodies is one that is being applied every day and all day long by children and savages. Unsophisticated humanity is constantly putting everything it sees up to its mouth in a frank spirit of experimental inquiry as to its gustatory properties. In civilised life we find everything ready labelled and assorted for us; we comparatively seldom require to roll the contents of a suspicious bottle (in very small quantities) doubtfully upon the tongue in order to discover whether it is pale sherry or Chili vinegar, Dublin stout or mushroom ketchup. But in the savage state, from which, geologically and biologically speaking, we have only just emerged, bottles and labels do not exist. Primitive man, therefore, in his sweet simplicity, has only two modes open before him for deciding whether the things he finds are or are not strictly edible. The first thing he does is to sniff at them; and smell, being, as Mr. Herbert Spencer has well put it, an anticipatory taste, generally gives him some idea of what the thing is likely to prove. The second thing he does is to pop it into his mouth, and proceed practically to examine its further characteristics.

Strictly speaking, with the tip of the tongue one can't really taste at all. If you put a small drop of honey or of oil of bitter almonds on that part of the mouth, you will find (no doubt to your great surprise) that it produces no effect of any sort; you only taste it when it begins slowly to diffuse itself, and reaches the true tasting region in the middle distance. But if you put a little cayenne or mustard on the same part, you will find that it bites you immediately—the experiment should be tried sparingly—while if you put it lower down in the mouth you will swallow it almost without noticing the pungency of the stimulant. The reason is, that the tip of the tongue is supplied only with nerves which are really nerves of touch, not nerves of taste proper; they belong to a totally different main branch, and they go to a different centre in the brain, together with the very similar threads which supply the nerves of smell for mustard and pepper. That is why the smell and taste of these pungent substances are so much alike, as everybody must have noticed, a good sniff at a mustard-pot producing almost the same irritating effects as an incautious mouthful. As a rule we don't accurately distinguish, it is true, between these different regions of taste in the mouth in ordinary life; but that is because we usually roll our food about instinctively, without paying much attention to the particular part affected by it. Indeed, when one is trying deliberate experiments in the subject, in order to test the varying sensitiveness of the different parts to different substances, it is necessary to keep the tongue quite dry, in order to isolate the thing you are experimenting with, and prevent its spreading to all parts of the mouth together. In actual practice this result is obtained in a rather ludicrous manner—by blowing upon the tongue, between each experiment, with a pair of bellows. To such undignified expedients does the pursuit of science lead the ardent modern psychologist. Those domestic rivals of Dr. Forbes Winslow, the servants, who behold the enthusiastic investigator alternately drying his tongue in this ridiculous fashion, as if he were a blacksmith's fire, and then squeezing out a single drop of essence of pepper, vinegar, or beef-tea from a glass syringe upon the dry surface, not unnaturally arrive at the conclusion that master has gone stark mad, and that, in their private opinion, it's the microscope and the skeleton as has done it.

Above all things, we don't want to be flayed alive. So the kinds of tastes discriminated by the tip of the tongue are the pungent, like pepper, cayenne and mustard; the astringent, like borax and alum; the alkaline, like soda and potash; the acid, like vinegar and green fruit; and the saline, like salt and ammonia. Almost all the bodies likely to give rise to such tastes (or, more correctly, sensations of touch in the tongue) are obviously unwholesome and destructive in their character, at least when taken in large quantities. Nobody wishes to drink nitric acid by the quart. The first business of this part of the tongue is, therefore, to warn us emphatically against caustic substances and corrosive acids, against vitriol and kerosene, spirits of wine and ether, capsicums and burning leaves or roots, such as those of the common English lords-and-ladies. Things of this sort are immediately destructive to the very tissues of the tongue and palate; if taken incautiously in too large doses, they burn the skin off the roof of the mouth; and when swallowed they play havoc, of course, with our internal arrangements. It is highly advisable, therefore, to have an immediate warning of these extremely dangerous substances, at the very outset of our feeding apparatus.

This kind of taste hardly differs from touch or burning. The sensibility of the tip of the tongue is only a very slight modification of the sensibility possessed by the skin generally, and especially by the inner folds over all delicate parts of the body. We all know that common caustic burns us wherever it touches; and it burns the tongue only in a somewhat more marked manner. Nitric or sulphuric acid attacks the fingers each after its own kind. A mustard plaster makes us tingle almost immediately; and the action of mustard on the tongue hardly differs, except in being more instantaneous and more discriminative. Cantharides work in just the same way. If you cut a red pepper in two and rub it on your neck, it will sting just as it does when put into soup (this experiment, however, is best tried upon one's younger brother; if made personally, it hardly repays the trouble and annoyance). Even vinegar and other acids, rubbed into the skin, are followed by a slight tingling; while the effect of brandy, applied, say, to the arms, is gently stimulating and pleasurable, somewhat in the same way as when normally swallowed in conjunction with the habitual seltzer. In short, most things which give rise to distinct tastes when applied to the tip of the tongue give rise to fainter sensations when applied to the skin generally. And one hardly needs to be reminded that pepper or vinegar placed (accidentally as a rule) on the inner surface of the eyelids produces a very distinct and unpleasant smart.

The fact is, the liability to be chemically affected by pungent or acid bodies is common to every part of the skin; but it is least felt where the tough outer skin is thickest, and most felt where that skin is thinnest, and the nerves are most plentifully distributed near the surface. A mustard plaster would probably fail to draw at all on one's heel or the palm of one's hand; while it is decidedly painful on one's neck or chest; and a mere speck of mustard inside the eyelid gives one positive torture for hours together. Now, the tip of the tongue is just a part of one's body specially set aside for this very object, provided with an extremely thin skin, and supplied with an immense number of nerves, on purpose so as to be easily affected by all such pungent, alkaline, or spirituous substances. Sir Wilfrid Lawson would probably conclude that it was deliberately designed by Providence to warn us against a wicked indulgence in the brandy and seltzer aforesaid.

At first sight it might seem as though there were hardly enough of such pungent and fiery things in existence to make it worth while for us to be provided with a special mechanism for guarding against them. That is true enough, no doubt, as regards our modern civilised life; though, even now, it is perhaps just as well that our children should have an internal monitor (other than conscience) to dissuade them immediately from indiscriminate indulgence in photographic chemicals, the contents of stray medicine bottles, and the best dried West India chilies. But in an earlier period of progress, and especially in tropical countries (where the Darwinians have now decided the human race made its first début upon this or any other stage), things were very different indeed. Pungent and poisonous plants and fruits abounded on every side. We have all of us in our youth been taken in by some too cruelly waggish companion, who insisted upon making us eat the bright, glossy leaves of the common English arum, which without look pretty and juicy enough, but within are full of the concentrated essence of pungency and profanity. Well, there are hundreds of such plants, even in cold climates, to tempt the eyes and poison the veins of unsuspecting cattle or childish humanity. There is buttercup, so horribly acrid that cows carefully avoid it in their closest cropped pastures; and yet your cow is not usually a too dainty animal. There is aconite, the deadly poison with which Dr. Lamson removed his troublesome relatives. There is baneberry, whose very name sufficiently describes its dangerous nature. There are horse-radish, and stinging rocket, and biting wall-pepper, and still smarter water-pepper, and worm-wood, and nightshade, and spurge, and hemlock, and half a dozen other equally unpleasant weeds. All of these have acquired their pungent and poisonous properties, just as nettles have acquired their sting, and thistles their thorns, in order to prevent animals from browsing upon them and destroying them. And the animals in turn have acquired a very delicate sense of pungency on purpose to warn them beforehand of the existence of such dangerous and undesirable qualities in the plants which they might otherwise be

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