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hard parts and could leave no evidence of their existence in the record of the rocks. And of their supposed descendants we find so few traces in the pre- Cambrian strata that the first steps in organic evolution must be supplied from such analogies in embryology as the following. The fertilized ovum, the cell with which each animal begins its life, grows and multiplies by cell division, and develops into a hollow globe of cells called the BLASTOSPHERE. This stage is succeeded by the stage of the GASTRULA,—an ovoid or cup-shaped body with a double wall of cells inclosing a body cavity, and with an opening, the primitive mouth. Each of these early embryological stages is represented by living animals,—the undivided cell by the PROTOZOA, the blastosphere by some rare forms, and the gastrula in the essential structure of the COELENTERATES,—the subkingdom to which the fresh-water hydra and the corals belong. All forms of animal life, from the coelenterates to the mammals, follow the same path in their embryological development as far as the gastrula stage, but here their paths widely diverge, those of each subkingdom going their own separate ways.

We may infer, therefore, that during the pre-Cambrian periods organic evolution followed the lines thus dimly traced. The earliest one-celled protozoa were probably succeeded by many- celled animals of the type of the blastosphere, and these by gastrula-like organisms. From the gastrula type the higher sub- divisions of animal life probably diverged, as separate branches from a common trunk. Much or all of this vast differentiation was accomplished before the opening of the next era; for all the subkingdoms are represented in the Cambrian except the vertebrates.

EVIDENCES OF PRE-CAMBRIAN LIFE. An indirect evidence of life during the pre-Cambrian periods is found in the abundant and varied fauna of the next period; for, if the theory of evolution is correct, the differentiation of the Cambrian fauna was a long process which might well have required for its accomplishment a large part of pre-Cambrian time.

Other indirect evidences are the pre-Cambrian limestones, iron ores, and graphite deposits, since such minerals and rocks have been formed in later times by the help of organisms. If the carbonate of lime of the Algonkian limestones and marbles was extracted from sea water by organisms, as is done at present by corals, mollusks, and other humble animals and plants, the life of those ancient seas must have been abundant. Graphite, a soft black mineral composed of carbon and used in the manufacture of lead pencils and as a lubricant, occurs widely in the metamorphic pre- Cambrian rocks. It is known to be produced in some cases by the metamorphism of coal, which itself is formed of decomposed vegetal tissues. Seams of graphite may therefore represent accumulations of vegetal matter such as seaweed. But limestone, iron ores, and graphite can be produced by chemical processes, and their presence in the pre-Cambrian makes it only probable, and not certain, that life existed at that time.

PRE-CAMBRIAN FOSSILS. Very rarely has any clear trace of an organism been found in the most ancient chapters of the geological record, so many of their leaves have been destroyed and so far have their pages been defaced. Omitting structures whose organic nature has been questioned, there are left to mention a tiny seashell of one of the most lowly types,—a DISCINA from the pre- Cambrian rocks of the Colorado Canyon,—and from the pre-Cambrian rocks of Montana trails of annelid worms and casts of their burrows in ancient beaches, and fragments of the tests of crustaceans. These diverse forms indicate that before the Algonkian had closed, life was abundant and had widely differentiated. We may expect that other forms will be discovered as the rocks are closely searched.

PRE-CAMBRIAN GEOGRAPHY. Our knowledge is far too meager to warrant an attempt to draw the varying outlines of sea and land during the Archean and Algonkian eras. Pre-Cambrian time probably was longer than all later geological time down to the present, as we may infer from the vast thicknesses of its rocks and the unconformities which part them. We know that during its long periods land masses again and again rose from the sea, were worn low, and were submerged and covered with the waste of other lands. But the formations of separated regions cannot be correlated because of the absence of fossils, and nothing more can be made out than the detached chapters of local histories, such as the outline given of the district about Lake Superior.

The pre-Cambrian rocks show no evidence of any forces then at work upon the earth except the forces which are at work upon it now. The most ancient sediments known are so like the sediments now being laid that we may infer that they were formed under conditions essentially similar to those of the present time. There is no proof that the sands of the pre-Cambrian sandstones were swept by any more powerful waves and currents than are offshore sands to-day, or that the muds of the pre-Cambrian shales settled to the sea floor in less quiet water than such muds settle in at present. The pre-Cambrian lands were, no doubt, worn by wind and weather, beaten by rain, and furrowed by streams as now, and, as now, they fronted the ocean with beaches on which waves dashed and along which tidal currents ran.

Perhaps the chief difference between the pre-Cambrian and the present was the absence of life upon the land. So far as we have any knowledge, no forests covered the mountain sides, no verdure carpeted the plains, and no animals lived on the ground or in the air. It is permitted to think of the most ancient lands as deserts of barren rock and rock waste swept by rains and trenched by powerful streams. We may therefore suppose that the processes of their destruction went on more rapidly than at present.

CHAPTER XVI THE CAMBRIAN

THE PALEOZOIC ERA. The second volume of the geological record, called the Paleozoic (Greek, PALAIOS, ancient; ZOE, life), has come down to us far less mutilated and defaced than has the first volume, which contains the traces of the most ancient life of the globe. Fossils are far more abundant in the Paleozoic than in the earlier strata, while the sediments in which they were entombed have suffered far less from metamorphism and other causes, and have been less widely buried from view, than the strata of the pre-Cambrian groups. By means of their fossils we can correlate the formations of widely separated regions from the beginning of the Paleozoic on, and can therefore trace some outline of the history of the continents.

Paleozoic time, although shorter than the pre-Cambrian as measured by the thickness of the strata, must still be reckoned in millions of years. During this vast reach of time the changes in organisms were very great. It is according to the successive stages in the advance of life that the Paleozoic formations are arranged in five systems,—the CAMBRIAN, the ORDOVICIAN, the SILURIAN, the DEVONIAN, and the CARBONIFEROUS. On the same basis the first three systems are grouped together as the older Paleozoic, because they alike are characterized by the dominance of the invertebrates; while the last two systems are united in the later Paleozoic, and are characterized, the one by the dominance of fishes, and the other by the appearance of amphibians and reptiles.

Each of these systems is world-wide in its distribution, and may be recognized on any continent by its own peculiar fauna. The names first given them in Great Britain have therefore come into general use, while their subdivisions, which often cannot be correlated in different countries and different regions, are usually given local names.

The first three systems were named from the fact that their strata are well displayed in Wales. The Cambrian carries the Roman name of Wales, and the Ordovician and Silurian the names of tribes of ancient Britons which inhabited the same country. The Devonian is named from the English county Devon, where its rocks were early studied. The Carboniferous was so called from the large amount of coal which it was found to contain in Great Britain and continental Europe.

THE CAMBRIAN

DISTRIBUTION OF STRATA. The Cambrian rocks outcrop in narrow belts about the pre-Cambrian areas of eastern Canada and the Lake Superior region, the Adirondacks and the Green Mountains. Strips of Cambrian formations occupy troughs in the pre-Cambrian rocks of New England and the maritime provinces of Canada; a long belt borders on the west the crystalline rocks of the Blue Ridge; and on the opposite side of the continent the Cambrian reappears in the mountains of the Great Basin and the Canadian Rockies. In the Mississippi valley it is exposed in small districts where uplift has permitted the stripping off of younger rocks. Although the areas of outcrop are small, we may infer that Cambrian rocks were widely deposited over the continent of North America.

PHYSICAL GEOGRAPHY. The Cambrian system of North America comprises three distinct series, the LOWER CAMBRIAN, the MIDDLE CAMBRIAN, and the UPPER CAMBRIAN, each of which is characterized by its own peculiar fauna. In sketching the outlines of the continent as it was at the beginning of the Paleozoic, it must be remembered that wherever the Lower Cambrian formations now are found was certainly then sea bottom, and wherever the Lower Cambrian are wanting, and the next formations rest directly on pre-Cambrian rocks, was probably then land.

EARLY CAMBRIAN GEOGRAPHY. In this way we know that at the opening of the Cambrian two long, narrow mediterranean seas stretched from north to south across the continent. The eastern sea extended from the Gulf of St. Lawrence down the Champlain-Hudson valley and thence along the western base of the Blue Ridge south at least to Alabama. The western sea stretched from the Canadian Rockies over the Great Basin and at least as far south as the Grand Canyon of the Colorado in Arizona.

Between these mediterraneans lay a great central land which included the pre-Cambrian U-shaped area of the Laurentian peneplain, and probably extended southward to the latitude of New Orleans. To the east lay a land which we may designate as APPALACHIA, whose western shore line was drawn along the site of the present Blue Ridge, but whose other limits are quite unknown. The land of Appalachia must have been large, for it furnished a great amount of waste during the entire Paleozoic era, and its eastern coast may possibly have lain even beyond the edge of the present continental shelf. On the western side of the continent a narrow land occupied the site of the Sierra Nevada Mountains.

Thus, even at the beginning of the Paleozoic, the continental plateau of North America had already been left by crustal movements in relief above the abysses of the great oceans on either side. The mediterraneans which lay upon it were shallow, as their sediments prove. They were EPICONTINENTAL SEAS; that is, they rested UPON (Greek, EPI) the submerged portion of the continental plateau. We have no proof that the deep ocean ever occupied any part of where North America now is.

The Middle and Upper Cambrian strata are found together with the Lower Cambrian over the area of both the eastern and the western mediterraneans, so that here the sea continued during the entire period. The sediments throughout are those of shoal water. Coarse cross-bedded sandstones record the action of strong shifting currents which spread coarse waste near shore and winnowed it of finer stuff. Frequent ripple marks on the bedding planes of the strata prove that the loose sands of the sea floor were near enough to the surface to be agitated by waves and tidal currents. Sun cracks show that often the outgoing tide exposed large muddy flats to the drying action of the sun. The fossils, also, of the strata are of kinds related to those which now live in shallow waters near the shore.

The sediments which gathered in the mediterranean seas were very thick, reaching in places the enormous depth of ten thousand feet. Hence the bottoms of these seas were sinking troughs, ever filling with waste from the adjacent land as fast as they subsided.

LATE CAMBRIAN GEOGRAPHY. The formations of the Middle and Upper Cambrian are found resting unconformably on the pre-Cambrian rocks from New York westward into Minnesota and at various points in the interior, as in Missouri and in Texas. Hence after earlier Cambrian time the central land subsided, with much the same effect as if the Mississippi valley were now to lower gradually, and the Gulf of Mexico to spread northward until it entered Lake Superior. The Cambrian seas transgressed the central land and strewed far and wide behind their advancing beaches the sediments of the later Cambrian upon an eroded surface of pre-Cambrian rocks.

The succession of the Cambrian formations in North America records many minor oscillations and varying conditions of physical geography; yet on the whole it tells of widening seas and lowering lands. Basal conglomerates and coarse sandstones

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